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a Department of Renewable Resources, 4-42 Earth Sciences Building, University of Alberta, Edmonton, AB, T5G 2E1 Canada
b Department of Botany, 505 Buller Building, University of Manitoba, Winnipeg, MB, R3T 2N2 Canada
c Syncrude Canada Ltd., Edmonton Research Centre, 9421-17 Avenue, Edmonton, AB, T6N 1H4 Canada
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